Spatiotemporal Patterns and Driving Factors of Soil Organic Carbon Content: A Case Study from a Montane Region
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A comprehensive grasp of the vertical distribution profiles of soil organic carbon (SOC) and its underlying governing mechanisms is fundamental to assessing terrestrial carbon stocks. Research efforts in mountain ecosystems, however, frequently fall short in providing depth-resolved analysis. This study investigated the spatial patterns and dominant drivers of SOC content across the full 0–200 cm soil profile in the Dabie Mountain Area (DMA), a subtropical montane region in central China. By utilizing high-resolution (250 m) SOC data and multi-source environmental variables for topography, climate, and soil properties, we employed hots pot analysis (Getis-Ord Gi*) and the geographical detector model to quantify spatial clustering and identify key influencing factors across six depth intervals in the DMA. The results showed that SOC content decreases exponentially with depth, with the surface 0–5 cm layer containing the highest concentration. A distinct shift in spatial organization occurred at an approximate depth of 60 cm: surface layers (0–60 cm) exhibited strong, clustered patterns (hot spots and cold spots), whereas deeper layers (>60 cm) transitioned to a more dispersed and spatially homogeneous distribution. Factor detection identified elevation and soil bulk density (SBD) as the most influential factors overall. More importantly, interaction detection revealed a depth-dependent transition in the dominant controlling complexes. In surface soils (0–15 cm), SOC heterogeneity was primarily governed by the interaction between topography (elevation) and soil properties (pH and SBD). In contrast, within the subsoil (15–200 cm), the interaction between climatic factors (temperature, precipitation) and soil properties (pH, SBD) became dominant. These findings demonstrated a fundamental shift from a topo-edaphic control regime in surface layers to a climate-edaphic control regime in deeper layers. This study provided a novel, three-dimensional perspective on SOC storage in mountains, highlighting the necessity of depth-resolved analyses for accurate carbon accounting and for formulating stratified land management strategies aimed at soil carbon conservation.
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[1] Almendros, G., & González-Pérez, J. A. (2025). Soil Organic Carbon Sequestration Mechanisms and the Chemical Nature of Soil Organic Matter—A Review. Sustainability (Switzerland), 17(15), 6689. doi:10.3390/su17156689.
[2] Scharlemann, J. P. W., Tanner, E. V. J., Hiederer, R., & Kapos, V. (2014). Global soil carbon: Understanding and managing the largest terrestrial carbon pool. Carbon Management, 5(1), 81–91. doi:10.4155/cmt.13.77.
[3] Qing, Z., Liu, H., Meng, X., Du, B., Zhang, S., & Yu, M. (2025). Assessment of the synergistic effects of future climate change and land use on soil organic carbon stock in Northeast China. Catena, 260, 109456. doi:10.1016/j.catena.2025.109456.
[4] Hari, M., & Tyagi, B. (2022). Terrestrial carbon cycle: tipping edge of climate change between the atmosphere and biosphere ecosystems. Environmental Science: Atmospheres, 2(5), 867–890. doi:10.1039/d1ea00102g.
[5] Zhang, Y., An, C. B., Zhang, W. S., Zheng, L. Y., Zhang, Y. Z., Lu, C., & Liu, L. Y. (2023). Drivers of mountain soil organic carbon stock dynamics: A review. Journal of Soils and Sediments, 23(1), 64-76. doi:10.1007/s11368-022-03313-w.
[6] Zhang, L., Xu, T., Bai, Y., Wiesmeier, M., Li, H., Huang, Y., Liu, Y., Xie, B., Song, M., Wu, J., & Liu, C. (2025). Historical and future dynamics of soil organic carbon and driving mechanisms in mountainous soils of China. Catena, 258, 109212. doi:10.1016/j.catena.2025.109212.
[7] Li, S., Zhang, A., Song, H., Guo, W., Tang, Z., Lei, G., & Qi, L. (2023). The Dominant Factor Affecting Soil Organic Carbon in Subtropical Phyllostachys edulis Forests Is Climatic Factors Rather Than Soil Physicochemical Properties. Forests, 14(5), 958. doi:10.3390/f14050958.
[8] Blackburn, K. W., Libohova, Z., Adhikari, K., Kome, C., Maness, X., & Silman, M. R. (2022). Influence of Land Use and Topographic Factors on Soil Organic Carbon Stocks and Their Spatial and Vertical Distribution. Remote Sensing, 14(12), 2846. doi:10.3390/rs14122846.
[9] Wang, G., Li, J., Mao, J., Fan, L., Ma, X., Zhang, W., Liang, Y., Hui, T., & Li, Y. (2025). Multiscale drivers and tipping points regulating particulate and mineral-associated organic carbon across Central Asian grasslands. Geoderma, 464, 117610. doi:10.1016/j.geoderma.2025.117610.
[10] Bai, R., Zhao, X., Wang, X., Lv, W., Li, J., Yang, F., Shangguan, Z., & Deng, L. (2026). SOC erosion reduction of the “Grain for green” program on the Loess Plateau, China. Soil and Tillage Research, 256, 106863. doi:10.1016/j.still.2025.106863.
[11] Guo, B., Fang, M., Yang, L., Guo, T., Ma, C., Hu, X., Guo, Z., Ma, Z., Li, Q., Wang, Z., & Liu, W. (2026). Remapping carbon storage change in retired farmlands on the Loess Plateau in China from 2000–2021 in high spatiotemporal resolution. Earth System Science Data, 18(1), 429–441. doi:10.5194/essd-18-429-2026.
[12] Tiruneh, G. A., Righi, C. A., Polizel, J. L., Gonçalves, V., & Pereira, C. R. (2026). Modeling soil organic carbon in the Brazilian amazon with geostatistical and machine learning techniques. Trees, Forests and People, 23. doi:10.1016/j.tfp.2026.101150.
[13] Wan, Q., Zhu, G., Guo, H., Zhang, Y., Pan, H., Yong, L., & Ma, H. (2019). Influence of Vegetation Coverage and Climate Environment on Soil Organic Carbon in the Qilian Mountains. Scientific Reports, 9(1), 17623. doi:10.1038/s41598-019-53837-4.
[14] Mishra, U., & Riley, W. J. (2015). Scaling impacts on environmental controls and spatial heterogeneity of soil organic carbon stocks. Biogeosciences, 12(13), 3993–4004. doi:10.5194/bg-12-3993-2015.
[15] Wiesmeier, M., Urbanski, L., Hobley, E., Lang, B., von Lützow, M., Marin-Spiotta, E., van Wesemael, B., Rabot, E., Ließ, M., Garcia-Franco, N., Wollschläger, U., Vogel, H. J., & Kögel-Knabner, I. (2019). Soil organic carbon storage as a key function of soils - A review of drivers and indicators at various scales. Geoderma, 333, 149–162. doi:10.1016/j.geoderma.2018.07.026.
[16] Hobley, E., Wilson, B., Wilkie, A., Gray, J., & Koen, T. (2015). Drivers of soil organic carbon storage and vertical distribution in Eastern Australia. Plant and Soil, 390(1–2), 111–127. doi:10.1007/s11104-015-2380-1.
[17] Zhou, T., Lv, Y., Xie, B., Xu, L., Zhou, Y., Mei, T., Li, Y., Yuan, N., & Shi, Y. (2023). Topography and Soil Organic Carbon in Subtropical Forests of China. Forests, 14(5), 1023. doi:10.3390/f14051023.
[18] Kuśmierz, S., Skowrońska, M., Tkaczyk, P., Lipiński, W., & Mielniczuk, J. (2023). Soil Organic Carbon and Mineral Nitrogen Contents in Soils as Affected by Their pH, Texture and Fertilization. Agronomy, 13(1), 267. doi:10.3390/agronomy13010267.
[19] Liu, Y., Chen, Y., Wu, Z., Wang, B., & Wang, S. (2021). Geographical detector-based stratified regression kriging strategy for mapping soil organic carbon with high spatial heterogeneity. Catena, 196, 104953. doi:10.1016/j.catena.2020.104953.
[20] Liu, Z., Lei, H., Sheng, H., & Wang, Y. (2023). Analysis of soil organic matter influencing factors in the Huangshui River Basin by using the optimal parameter-based geographical detector model. Geocarto International, 38(1), 2246935. doi:10.1080/10106049.2023.2246935.
[21] Dubeux Jr, J. C., Lira Junior, M. D. A., Simili, F. F., Bretas, I. L., Trumpp, K. R., Bizzuti, B. E., Garcia, L., Oduor, K. T., Queiroz, L. M. D., Acuña, J. P., & Mendes, C. T. (2024). Deep soil organic carbon: A review. CABI Reviews, 2024(2), 1-17. doi:10.1079/cabireviews.2024.002.
[22] Zeng, X. M., Bastida, F., Plaza, C., Zhou, G., Vera, A., Liu, Y. R., & Delgado-Baquerizo, M. (2023). The Contribution of Biotic Factors in Explaining the Global Distribution of Inorganic Carbon in Surface Soils. Global Biogeochemical Cycles, 37(10), 7957. doi:10.1029/2023GB007957.
[23] Song, B., Wang, M., Zhang, S., Zhang, L., Lu, Y., Guo, H., Guo, X., Zhang, Y., & Zhou, X. (2025). Spatial distribution, drivers, and future variation of soil organic carbon in China’s ecosystems: A meta-analysis and machine-learning assessment. Ecological Indicators, 179, 114255. doi:10.1016/j.ecolind.2025.114255.
[24] Ghosh, A., Singh, A. K., Kumar, S., Manna, M. C., Bhattacharyya, R., Agnihortri, R., Singh Gahlaud, S. K., Sannagoudar, M. S., Gautam, K., Kumar, R. V., & Chaudhari, S. K. (2020). Differentiating biological and chemical factors of top and deep soil carbon sequestration in semi-arid tropical Inceptisol: an outcome of structural equation modeling. Carbon Management, 11(5), 441–453. doi:10.1080/17583004.2020.1796143.
[25] Nie, X., Wang, D., Yang, L., & Zhou, G. (2021). Controls on variation of soil organic carbon concentration in the shrublands of the north-eastern Tibetan Plateau. European Journal of Soil Science, 72(4), 1817–1830. doi:10.1111/ejss.13084.
[26] Zhang, R., Liu, X., Heathman, G. C., Yao, X., Hu, X., & Zhang, G. (2013). Assessment of soil erosion sensitivity and analysis of sensitivity factors in the Tongbai-Dabie mountainous area of China. Catena, 101, 92–98. doi:10.1016/j.catena.2012.10.008.
[27] Yang, S. Y., Yang, K. F., Zhang, X. T., & Wang, J. (2011). The research of forest soil organic carbon accumulation in Dabie Mountain. Meteorological and Environmental Research, (3), 43-46. doi:CNKI:SUN:MEVR.0.2011-03-014.
[28] Qin, J., Liu, Y., Bi, Q., Chen, Z., & Zhang, B. (2023). Response of leaf and soil C, N and P stoichiometry in different Pinus massoniana forest types to slope aspect in the Dabie mountains region of North subtropical, China. Frontiers in Environmental Science, 11, 1148986. doi:10.3389/fenvs.2023.1148986.
[29] Fang, L., Liu, Y., Li, C., & Cai, J. (2023). Spatiotemporal Characteristics and Future Scenario Simulation of the Trade-offs and Synergies of Mountain Ecosystem Services: A Case Study of the Dabie Mountains Area, China. Chinese Geographical Science, 33(1), 144–160. doi:10.1007/s11769-023-1330-8.
[30] Dong, Q., Song, C., Wen, H., Xiang, J., Wang, P., & Yan, M. (2024). Comprehensive Geochemical Evaluation and Influencing Factors of Topsoil Nutrients in a Farming Area of Dabie Mountain in Western Anhui, China. Yankuang Ceshi, 43(2), 344–355. doi:10.15898/j.ykcs.202206180117.
[31] Zheng, L., Sun, J., Qiu, X., & Yang, Z. (2020). Five-year climatology of local convections in the Dabie mountains. Atmosphere, 11(11), 1246. doi:10.3390/atmos11111246.
[32] Hou, Y., & Dai, Y. (2024). Spatial Configuration and Sustainable Conservation of Ecotourism Resources in the Dabie Mountains, Eastern China, Using an Ecosystem Services Model. Diversity, 16(12), 782. doi:10.3390/d16120782.
[33] Qin, X., Li, J., & Li, N. (2025). Fine-Scale Spatiotemporal Distribution Characteristics of Precipitation in the Dabie Mountains Region. Journal of Applied Meteorology and Climatology, 64(12), 1803–1817. doi:10.1175/JAMC-D-25-0014.1.
[34] Yang, T., Wu, F., Luo, M., Xiong, J., Nie, X., Cao, F., Ruan, Y., Li, F., Huang, W., Liang, T., & Yang, Y. (2024). Accumulation Pattern and Potential Ecological Risk of Heavy Metals in Topsoil as Affected by Diverse Sources in Different Ecosystems in Western Dabie Mountain. Forests, 15(7), 1116. doi:10.3390/f15071116.
[35] Li, G., Yang, T., Chen, R., Dong, H., Wu, F., Zhan, Q., Huang, J., Luo, M., & Wang, L. (2025). Experimental study on in-situ simulation of rainfall-induced soil erosion in forest lands converted to cash crop areas in Dabie Mountains. PLoS ONE, 20(2 February), 317889. doi:10.1371/journal.pone.0317889.
[36] Zhu, Y., Du, C., Sun, L., Liu, X., Jamshidi, A. H., & Zhang, S. (2024). Impacts of intercropping tea trees under forest on soil water infiltration in the northern of Dabie Mountains. Nongye Gongcheng Xuebao/Transactions of the Chinese Society of Agricultural Engineering, 40(19), 72–82. doi:10.11975/j.issn.1002-6819.202403063.
[37] Zhu, Y., Sun, L., Jamshidi, A. H., Liu, X., Zheng, Y., & Fan, Z. (2025). Effect of forest conversion on soil water infiltration in the Dabie mountainous area, China. Journal of Hydrology: Regional Studies, 59, 102351. doi:10.1016/j.ejrh.2025.102351.
[38] Loess Plateau SubCenter (2026). National Earth System Science Data Center, National Science & Technology Infrastructure of China. Available online: http://loess.geodata.cn (accessed on May 2026)
[39] Karra, K., Kontgis, C., Statman-Weil, Z., Mazzariello, J. C., Mathis, M., & Brumby, S. P. (2021). Global Land Use/Land Cover with Sentinel 2 and Deep Learning. International Geoscience and Remote Sensing Symposium (IGARSS), 2021-July, 4704–4707. doi:10.1109/IGARSS47720.2021.9553499.
[40] Xie, W.-F., Li, J.-K., Peng, K., Zhang, K., & Ullah, Z. (2024). The Application of Local Moran’s I and Getis–Ord Gi* to Identify Spatial Patterns and Critical Source Areas of Agricultural Nonpoint Source Pollution. Journal of Environmental Engineering, 150(5), 4024011. doi:10.1061/joeedu.eeeng-7585.
[41] Boubekraoui, H., Maouni, Y., Ghallab, A., Draoui, M., & Maouni, A. (2023). Spatio-temporal analysis and identification of deforestation hotspots in the Moroccan western Rif. Trees, Forests and People, 12, 100388. doi:10.1016/j.tfp.2023.100388.
[42] Nie, Q., Wu, G., Li, L., Man, W., Ma, J., Bao, Z., Luo, L., & Li, H. (2024). Exploring scaling differences and spatial heterogeneity in drivers of carbon storage Changes: A comprehensive geographic analysis framework. Ecological Indicators, 165, 112193. doi:10.1016/j.ecolind.2024.112193.
[43] Wang, J. F., Li, X. H., Christakos, G., Liao, Y. L., Zhang, T., Gu, X., & Zheng, X. Y. (2010). Geographical detectors-based health risk assessment and its application in the neural tube defects study of the Heshun Region, China. International Journal of Geographical Information Science, 24(1), 107–127. doi:10.1080/13658810802443457.
[44] Wang, J. F., Zhang, T. L., & Fu, B. J. (2016). A measure of spatial stratified heterogeneity. Ecological Indicators, 67, 250–256. doi:10.1016/j.ecolind.2016.02.052.
[45] Wu, Z., Liu, Y., Li, G., Han, Y., Li, X., & Chen, Y. (2022). Influences of Environmental Variables and Their Interactions on Chinese Farmland Soil Organic Carbon Density and Its Dynamics. Land, 11(2), 208. doi:10.3390/land11020208.
[46] Chen, L., & Shi, L. (2024). Differences in urban–rural gradient and driving factors of PM2.5 concentration in the Zhengzhou Metropolitan Area. Air Quality, Atmosphere and Health, 17(10), 2187–2201. doi:10.1007/s11869-024-01564-9.
[47] Wang, J., & Xu, C. (2017). Geodetector: Principle and prospective. Dili Xuebao/Acta Geographica Sinica, 72(1), 116–134. doi:10.11821/dlxb201701010.
[48] Song, Y., Wang, J., Ge, Y., & Xu, C. (2020). An optimal parameters-based geographical detector model enhances geographic characteristics of explanatory variables for spatial heterogeneity analysis: cases with different types of spatial data. GIScience and Remote Sensing, 57(5), 593–610. doi:10.1080/15481603.2020.1760434.
[49] Wu, J. (2004). Effects of changing scale on landscape pattern analysis: Scaling relations. Landscape Ecology, 19(2), 125–138. doi:10.1023/B:LAND.0000021711.40074.ae.
[50] Arrouays, D., McBratney, A. B., Minasny, B., Hempel, J. W., Heuvelink, G. B. M., McMillan, R. A., Hartemink, A. E., Lagacherie, P., & McKenzie, N. J. (2013). GlobalSoilMap. Basis for the global spatial soil information system. CRC Press, Florida, United States.
[51] Li, A. W., Ran, M., Song, L. Y., Xue, J. L., Zhang, Y. Y., Li, C. J., Deng, Q., Fang, H. Y., Dai, T. F., & Li, Q. Q. (2023). Spatial Distribution Characteristics and Influencing Factors of Cropland Topsoil Organic Carbon Content in the Sichuan Basin. Resources and Environment in the Yangtze Basin, 32(5), 1102–1112. doi:10.11870/cjlyzyyhj202305019.
[52] Jobbágy, E. G., & Jackson, R. B. (2000). The vertical distribution of soil organic carbon and its relation to climate and vegetation. Ecological Applications, 10(2), 423–436. doi:10.1890/1051-0761(2000)010[0423:TVDOSO]2.0.CO;2.
[53] Jackson, R. B., Lajtha, K., Crow, S. E., Hugelius, G., Kramer, M. G., & Piñeiro, G. (2017). The Ecology of Soil Carbon: Pools, Vulnerabilities, and Biotic and Abiotic Controls. Annual Review of Ecology, Evolution, and Systematics, 48(1), 419–445. doi:10.1146/annurev-ecolsys-112414-054234.
[54] Rumpel, C., & Kögel-Knabner, I. (2011). Deep soil organic matter-a key but poorly understood component of terrestrial C cycle. Plant and Soil, 338(1), 143–158. doi:10.1007/s11104-010-0391-5.
[55] Egli, M., Mirabella, A., & Sartori, G. (2008). The role of climate and vegetation in weathering and clay mineral formation in late Quaternary soils of the Swiss and Italian Alps. Geomorphology, 102(3–4), 307–324. doi:10.1016/j.geomorph.2008.04.001.
[56] Gami, S. K., Lauren, J. G., & Duxbury, J. M. (2009). Soil organic carbon and nitrogen stocks in Nepal long-term soil fertility experiments. Soil and Tillage Research, 106(1), 95–103. doi:10.1016/j.still.2009.10.003.
[57] Nielsen, D. R., & Bouma, J. (1985). Soil spatial variability: Proceedings of a workshop of the ISSS and the SSSA, Las Vegas, USA (Pudoc 296). Centre for Agricultural Publishing and Documentation, United States.
[58] Wilding, L. P. (1985). Spatial variability: its documentation, accommodation and implication to soil surveys. Soil Spatial Variability, 166–189.
[59] Schmidt, M. W. I., Torn, M. S., Abiven, S., Dittmar, T., Guggenberger, G., Janssens, I. A., Kleber, M., Kögel-Knabner, I., Lehmann, J., Manning, D. A. C., Nannipieri, P., Rasse, D. P., Weiner, S., & Trumbore, S. E. (2011). Persistence of soil organic matter as an ecosystem property. Nature, 478(7367), 49–56. doi:10.1038/nature10386.
[60] Grüneberg, E., Schöning, I., Hessenmöller, D., Schulze, E. D., & Weisser, W. W. (2013). Organic layer and clay content control soil organic carbon stocks in density fractions of differently managed German beech forests. Forest Ecology and Management, 303, 1–10. doi:10.1016/j.foreco.2013.03.014.
[61] Niu, X., Liu, C., Jia, X., & Zhu, J. (2021). Changing soil organic carbon with land use and management practices in a thousand-year cultivation region. Agriculture, Ecosystems and Environment, 322, 107639. doi:10.1016/j.agee.2021.107639.
[62] Yang, X., Xu, J., Wang, H., Quan, H., Yu, H., Luan, J., Wang, D., Li, Y., & Lv, D. (2024). Vertical distribution characteristics of soil organic carbon and vegetation types under different elevation gradients in Cangshan, Dali. PeerJ, 12, 16686. doi:10.7717/peerj.16686.
[63] Batjes, N. H. (2014). Total carbon and nitrogen in the soils of the world. European Journal of Soil Science, 65(1), 10–21. doi:10.1111/ejss.12114_2.
[64] Mishra, U., Lal, R., Slater, B., Calhoun, F., Liu, D., & Van Meirvenne, M. (2009). Predicting Soil Organic Carbon Stock Using Profile Depth Distribution Functions and Ordinary Kriging. Soil Science Society of America Journal, 73(2), 614–621. doi:10.2136/sssaj2007.0410.
[65] Zhang, Z., Huang, X., & Zhou, Y. (2020). Spatial heterogeneity of soil organic carbon in a karst region under different land use patterns. Ecosphere, 11(3), 3077. doi:10.1002/ecs2.3077.
[66] Zhou, J., Wang, Y., Tong, Y., Sun, H., Zhao, Y., & Zhang, P. (2023). Regional spatial variability of soil organic carbon in 0–5 m depth and its dominant factors. Catena, 231, 107326. doi:10.1016/j.catena.2023.107326.
[67] Deng, L., Liu, G. bin, & Shangguan, Z. ping. (2014). Land-use conversion and changing soil carbon stocks in China’s “Grain-for-Green” Program: A synthesis. Global Change Biology, 20(11), 3544–3556. doi:10.1111/gcb.12508.
[68] Conant, R. T., Cerri, C. E. P., Osborne, B. B., & Paustian, K. (2017). Grassland management impacts on soil carbon stocks: A new synthesis: A. Ecological Applications, 27(2), 662–668. doi:10.1002/eap.1473.
[69] Kayranli, B., Scholz, M., Mustafa, A., & Hedmark, Å. (2010). Carbon storage and fluxes within freshwater wetlands: A critical review. Wetlands, 30(1), 111–124. doi:10.1007/s13157-009-0003-4.
[70] West, T. O., & Post, W. M. (2002). Soil Organic Carbon Sequestration Rates by Tillage and Crop Rotation. Soil Science Society of America Journal, 66(6), 1930–1946. doi:10.2136/sssaj2002.1930.
[71] Ogle, S. M., Breidt, F. J., & Paustian, K. (2005). Agricultural management impacts on soil organic carbon storage under moist and dry climatic conditions of temperate and tropical regions. Biogeochemistry, 72(1), 87–121. doi:10.1007/s10533-004-0360-2.
[72] Tsozué, D., Nghonda, J. P., Tematio, P., & Basga, S. D. (2019). Changes in soil properties and soil organic carbon stocks along an elevation gradient at Mount Bambouto, Central Africa. Catena, 175, 251–262. doi:10.1016/j.catena.2018.12.028.
[73] Zhang, Y., Ai, J., Sun, Q., Li, Z., Hou, L., Song, L., Tang, G., Li, L., & Shao, G. (2021). Soil organic carbon and total nitrogen stocks as affected by vegetation types and altitude across the mountainous regions in the Yunnan Province, south-western China. Catena, 196, 104872. doi:10.1016/j.catena.2020.104872.
[74] Phillips, J., Ramirez, S., Wayson, C., & Duque, A. (2019). Differences in carbon stocks along an elevational gradient in tropical mountain forests of Colombia. Biotropica, 51(4), 490–499. doi:10.1111/btp.12675.
[75] Girardin, C. A. J., Malhi, Y., Aragão, L. E. O. C., Mamani, M., Huaraca Huasco, W., Durand, L., Feeley, K. J., Rapp, J., Silva-Espejo, J. E., Silman, M., Salinas, N., & Whittaker, R. J. (2010). Net primary productivity allocation and cycling of carbon along a tropical forest elevational transect in the Peruvian Andes. Global Change Biology, 16(12), 3176–3192. doi:10.1111/j.1365-2486.2010.02235.x.
[76] He, P., Lu, J., Ren, Y., Li, J., Hou, L., Deng, X., Gao, T., & Cheng, F. (2023). Altitude and slope aspects as the key factors affecting the change of C:N:P stoichiometry in the leaf-litter-soil system of alpine timberline ecotones of the Sygera Mountains in Southeast Tibet, China. Geoderma Regional, 32, 602. doi:10.1016/j.geodrs.2022.e00602.
[77] Zhang, X., Li, X., Ji, X., Zhang, Z., Zhang, H., Zha, T., & Jiang, L. (2021). Elevation and total nitrogen are the critical factors that control the spatial distribution of soil organic carbon content in the shrubland on the Bashang Plateau, China. Catena, 204, 105415. doi:10.1016/j.catena.2021.105415.
[78] Song, X., Cao, M., Li, J., Kitching, R. L., Nakamura, A., Laidlaw, M. J., Tang, Y., Sun, Z., Zhang, W., & Yang, J. (2021). Different environmental factors drive tree species diversity along elevation gradients in three climatic zones in Yunnan, southern China. Plant Diversity, 43(6), 433–443. doi:10.1016/j.pld.2021.04.006.
[79] Xing, W., Lu, X., Ying, J., Lan, Z., Chen, D., & Bai, Y. (2022). Disentangling the effects of nitrogen availability and soil acidification on microbial taxa and soil carbon dynamics in natural grasslands. Soil Biology and Biochemistry, 164, 108495. doi:10.1016/j.soilbio.2021.108495.
[80] Sollins, P., & Gregg, J. W. (2017). Soil organic matter accumulation in relation to changing soil volume, mass, and structure: Concepts and calculations. Geoderma, 301, 60–71. doi:10.1016/j.geoderma.2017.04.013.
[81] Davidson, E. A., Trumbore, S. E., & Amundson, R. (2000). Soil warming and organic carbon content. Nature, 408(6814), 789–790. doi:10.1038/35048672.
[82] Chen, Q., Niu, B., Hu, Y., Luo, T., & Zhang, G. (2020). Warming and increased precipitation indirectly affect the composition and turnover of labile-fraction soil organic matter by directly affecting vegetation and microorganisms. Science of the Total Environment, 714, 136787. doi:10.1016/j.scitotenv.2020.136787.
[83] Zhang, L., Zheng, Q., Liu, Y., Liu, S., Yu, D., Shi, X., Xing, S., Chen, H., & Fan, X. (2019). Combined effects of temperature and precipitation on soil organic carbon changes in the uplands of eastern China. Geoderma, 337, 1105–1115. doi:10.1016/j.geoderma.2018.11.026.
[84] Zhuo, Z., Chen, Q., Zhang, X., Chen, S., Gou, Y., Sun, Z., Huang, Y., & Shi, Z. (2022). Soil organic carbon storage, distribution, and influencing factors at different depths in the dryland farming regions of Northeast and North China. Catena, 210, 105934. doi:10.1016/j.catena.2021.105934.
[85] Sun, T., Wang, Y., Hui, D., Jing, X., & Feng, W. (2020). Soil properties rather than climate and ecosystem type control the vertical variations of soil organic carbon, microbial carbon, and microbial quotient. Soil Biology and Biochemistry, 148, 107905. doi:10.1016/j.soilbio.2020.107905.
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